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Open Access

Peer-reviewed

Research Article

A Comparison of Facial Color Pattern and Gazing Behavior in Canid Species Suggests Gaze Communication in Gray Wolves (Canis lupus)

  • Sayoko Ueda ,

    sykueda.ookami@gmail.com

    Affiliations Tokyo Institute of Technology, Tokyo, Japan, Wildlife Research Center of Kyoto University, Kyoto, Japan

    ⨯
  • Gaku Kumagai,

    Affiliation Tama Zoological Park, Tokyo Zoological Park Society, Tokyo, Japan

    ⨯
  • Yusuke Otaki,

    Current address: Nogeyama Zoological Gardens, Yokohama, Japan

    Affiliation Yokohama Zoological Gardens, Yokohama, Japan

    ⨯
  • Shinya Yamaguchi,

    Current address: Nogeyama Zoological Gardens, Yokohama, Japan

    Affiliation Yokohama Zoological Gardens, Yokohama, Japan

    ⨯
  • Shiro Kohshima

    Affiliation Wildlife Research Center of Kyoto University, Kyoto, Japan

    ⨯

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