About the Authors

Monica A. Daley

mdaley@rvc.ac.uk

Affiliation Department of Comparative Biomedical Sciences, Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Dennis M. Bramble

Affiliation Department of Biology, University of Utah, Salt Lake City, Utah, United States of America

David R. Carrier

Affiliation Department of Biology, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors declare that co­author David Carrier is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MAD DRC DMB. Performed the experiments: MAD. Analyzed the data: MAD. Wrote the paper: MAD. Interpreted the data and revised the manuscript: DMB DRC.