About the Authors

Daniel H. Janzen

djanzen@sas.upenn.edu

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Winnie Hallwachs

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

John M. Burns

Affiliation Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, D.C., United States of America

Mehrdad Hajibabaei

Affiliation Department of Integrative Biology, Biodiversity Institute of Ontario, University of Guelph, Guelph, Canada

Claudia Bertrand

Affiliation Department of Integrative Biology, Biodiversity Institute of Ontario, University of Guelph, Guelph, Canada

Paul D. N. Hebert

Affiliation Department of Integrative Biology, Biodiversity Institute of Ontario, University of Guelph, Guelph, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DHJ WH JMB MH CB PDNH. Performed the experiments: DHJ WH JMB MH CB PDNH. Analyzed the data: DHJ WH JMB MH CB PDNH. Contributed reagents/materials/analysis tools: DHJ WH JMB MH CB PDNH. Wrote the paper: DHJ WH JMB MH CB PDNH.