About the Authors

Saori Tsuji

Contributed equally to this work with: Saori Tsuji, Fumihiko Kawamura

Affiliation Bio Frontier Project Promotion Section, Organization for Tottori Industrial Promotion, Yonago, Japan

Fumihiko Kawamura

Contributed equally to this work with: Saori Tsuji, Fumihiko Kawamura

Affiliation Division of Molecular Genetics and Biofunction, Department of Biomedical Science, Institute of Regenerative Medicine and Biofunction, Graduate School of Medical Science, Tottori University, Yonago, Japan

Musashi Kubiura

Affiliation Division of Molecular Genetics and Biofunction, Department of Biomedical Science, Institute of Regenerative Medicine and Biofunction, Graduate School of Medical Science, Tottori University, Yonago, Japan

Ayaka Hayashi

Affiliation Division of Molecular Genetics and Biofunction, Department of Biomedical Science, Institute of Regenerative Medicine and Biofunction, Graduate School of Medical Science, Tottori University, Yonago, Japan

Tetsuya Ohbayashi

Affiliation Division of Laboratory Animal Science, Research Center for Bioscience and Technology, Tottori University, Yonago, Japan

Yasuhiro Kazuki

Affiliations Division of Molecular Genetics and Biofunction, Department of Biomedical Science, Institute of Regenerative Medicine and Biofunction, Graduate School of Medical Science, Tottori University, Yonago, Japan, Chromosome Engineering Research Center, Tottori University, Yonago, Japan

Christophe Chesné

Affiliation Biopredic International, Rennes, France

Mitsuo Oshimura

Affiliation Chromosome Engineering Research Center, Tottori University, Yonago, Japan

Masako Tada

matada@med.tottori-u.ac.jp

Affiliation Chromosome Engineering Research Center, Tottori University, Yonago, Japan

Competing Interests

ST is an employee of Bio Frontier Project Promotion Section, Organization for Tottori Industrial Promotion, which holds no patent and production in development or marketed product to declare. This does not alter the authors' adherence to all the PLOS ONE policies on the sharing data and materials.

Author Contributions

Conceived and designed the experiments: MT. Performed the experiments: ST FK MT MK AH. Analyzed the data: ST MT. Contributed reagents/materials/analysis tools: TO YK CC MO. Wrote the paper: MT ST.