About the Authors

Carlo Ricotta

carlo.ricotta@uniroma1.it

Affiliation Department of Environmental Biology, University of Rome ‘La Sapienza’, Rome, Italy

Giovanni Bacaro

Affiliations CNR, Istituto di Ricerca per la Protezione Idrogeologica, Perugia, Italy, BIOCONNET, Biodiversity and Conservation Network, Department of Life Science, University of Siena, Siena, Italy

Marco Moretti

Affiliation Swiss Federal Research Institute WSL, Birmensdorf, Switzerland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CR. Analyzed the data: CR GB MM. Wrote the paper: CR GB MM. Collected the data: MM. Designed the software used in analysis: GB.