About the Authors

Carly Muletz

cmuletz@umd.edu

Affiliations Department of Biology, University of Maryland, College Park, Maryland, United States of America, Center for Conservation and Evolutionary Genetics, Smithsonian Conservation Biology Institute, National Zoological Park, Washington, DC, United States of America

Nicholas M. Caruso

Affiliations Department of Biology, University of Maryland, College Park, Maryland, United States of America, Department of Biological Sciences, University of Alabama, Tuscaloosa, Alabama, United States of America

Robert C. Fleischer

Affiliation Center for Conservation and Evolutionary Genetics, Smithsonian Conservation Biology Institute, National Zoological Park, Washington, DC, United States of America

Roy W. McDiarmid

Affiliation USGS Patuxent Wildlife Research Center, Smithsonian Institution, National Museum of Natural History, Washington, DC, United States of America

Karen R. Lips

Affiliation Department of Biology, University of Maryland, College Park, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CM NMC RCF RWM KRL. Performed the experiments: CM NMC. Analyzed the data: CM KRL. Contributed reagents/materials/analysis tools: RCF RWM KRL. Contributed to the writing of the manuscript: CM NMC RCF RWM KRL. Granted permission to use collections at the National Museum of Natural History for sampling specimens: RWM.