About the Authors

Richard T. McCrea

rtmccrea@prprc.com

Affiliations Peace Region Palaeontology Research Centre, Tumbler Ridge, British Columbia, Canada, Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta, Canada

Lisa G. Buckley

Affiliations Peace Region Palaeontology Research Centre, Tumbler Ridge, British Columbia, Canada, Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

James O. Farlow

Affiliation Department of Geosciences, Indiana-Purdue University, Fort Wayne, Indiana, United States of America

Martin G. Lockley

Affiliation Dinosaur Trackers Research Group, University of Colorado at Denver, Denver, Colorado, United States of America

Philip J. Currie

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

Neffra A. Matthews

Affiliation Bureau of Land Management, Denver, Colorado, United States of America

S. George Pemberton

Affiliation Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RTM LGB. Performed the experiments: RTM LGB NAM. Analyzed the data: RTM LGB JOF MGL PJC SGP. Contributed reagents/materials/analysis tools: RTM LGB JOF MGL PJC NAM. Contributed to the writing of the manuscript: RTM LGB JOF MGL PJC NAM SGP. Collected measured specimen data: RTM LGB.