About the Authors

John M. Carethers

jcarethe@umich.edu

Affiliations Division of Gastroenterology, Department of Internal Medicine, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Medicine, University of California San Diego, San Diego, California, United States of America, Moores Cancer Center, University of California San Diego, San Diego, California, United States of America

Bhavya Murali

Contributed equally to this work with: Bhavya Murali, Bing Yang, Ryan T. Doctolero, Akihiro Tajima, Kathleen L. McGuire

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Bing Yang

Contributed equally to this work with: Bhavya Murali, Bing Yang, Ryan T. Doctolero, Akihiro Tajima, Kathleen L. McGuire

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Ryan T. Doctolero

Contributed equally to this work with: Bhavya Murali, Bing Yang, Ryan T. Doctolero, Akihiro Tajima, Kathleen L. McGuire

Affiliation Department of Medicine, University of California San Diego, San Diego, California, United States of America

Akihiro Tajima

Contributed equally to this work with: Bhavya Murali, Bing Yang, Ryan T. Doctolero, Akihiro Tajima, Kathleen L. McGuire

Affiliation Department of Medicine, University of California San Diego, San Diego, California, United States of America

Ranor Basa

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

E. Julieta Smith

Affiliation Department of Medicine, University of California San Diego, San Diego, California, United States of America

Monte Lee

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Ryan Janke

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Tina Ngo

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Ruth Tejada

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Ming Ji

Affiliation Graduate School Public Health, San Diego State University, San Diego, California, United States of America

Matthew Kinseth

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Betty L. Cabrera

Affiliation Department of Medicine, University of California San Diego, San Diego, California, United States of America

Katsumi Miyai

Affiliation Department of Pathology, University of California San Diego, San Diego, California, United States of America

Temitope O. Keku

Affiliation Department of Medicine, University of North Carolina, Chapel Hill, North Carolina, United States of America

Christopher F. Martin

Affiliation Department of Medicine, University of North Carolina, Chapel Hill, North Carolina, United States of America

Joseph A. Galanko

Affiliation Department of Epidemiology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Robert S. Sandler

Affiliations Department of Medicine, University of North Carolina, Chapel Hill, North Carolina, United States of America, Department of Epidemiology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Kathleen L. McGuire

Contributed equally to this work with: Bhavya Murali, Bing Yang, Ryan T. Doctolero, Akihiro Tajima, Kathleen L. McGuire

Affiliation Department of Biology, Molecular Biology Institute, San Diego State University, San Diego, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JMC KLM. Performed the experiments: BM BY RTD AT RB EJS ML RJ TN RT MK BC. Analyzed the data: JMC JAG MJ KM KLM. Contributed reagents/materials/analysis tools: JMC TOK CFM RSS KLM. Wrote the paper: JMC KLM.