About the Authors

Hannah Grist

Affiliations Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen, Aberdeenshire, United Kingdom, Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom, Scottish Ornithologists' Club, Aberlady, East Lothian, United Kingdom

Francis Daunt

frada@ceh.ac.uk (FD); jane.reid@abdn.ac.uk (JMR)

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Sarah Wanless

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Emily J. Nelson

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Mike P. Harris

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Mark Newell

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Sarah Burthe

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Jane M. Reid

frada@ceh.ac.uk (FD); jane.reid@abdn.ac.uk (JMR)

Affiliation Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen, Aberdeenshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HG FD SW JR. Performed the experiments: HG FD SW EN MPH MN SB JR. Analyzed the data: HG FD SW JR. Wrote the paper: HG FD SW EN MPH MN SB JR.