About the Authors

Kewei Zhang

zhangkw@sdu.edu.cn

Affiliation School of Life Sciences, Shandong University, Ministry of Education Key Laboratory of Plant Cell Engineering and Germplasm Enhancement, Jinan, China

Hanhan Liu

Affiliation School of Life Sciences, Shandong University, Ministry of Education Key Laboratory of Plant Cell Engineering and Germplasm Enhancement, Jinan, China

Peilin Tao

Affiliation College of Agriculture Vocational, Xuzhou Biology Engineering Technical College, Xuzhou, China

Huan Chen

Affiliation School of Life Sciences, Shandong University, Ministry of Education Key Laboratory of Plant Cell Engineering and Germplasm Enhancement, Jinan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KZ. Performed the experiments: HL PT HC. Analyzed the data: KZ HL PT. Contributed reagents/materials/analysis tools: PT HL. Wrote the paper: KZ HL.