About the Authors

Nao Sugiyama

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Asako Murayama

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Ryosuke Suzuki

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Noriyuki Watanabe

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Masaaki Shiina

Affiliation Department of Gastroenterology and Hepatology, Shin-Yurigaoka General Hospital, Kawasaki, Kanagawa, Japan

T. Jake Liang

Affiliation Liver Diseases Branch, National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Takaji Wakita

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Takanobu Kato

takato@nih.go.jp

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TJL TW TK. Performed the experiments: NS AM RS NW MS TK. Analyzed the data: NS AM TK. Wrote the paper: NS AM MS TK.