About the Authors

Amy E. Koid

Affiliation Department of Biological Sciences, University of Southern California Los Angeles, Los Angeles, California, United States of America

Zhenfeng Liu

Affiliation Department of Biological Sciences, University of Southern California Los Angeles, Los Angeles, California, United States of America

Ramon Terrado

Affiliation Department of Biological Sciences, University of Southern California Los Angeles, Los Angeles, California, United States of America

Adriane C. Jones

Affiliation Department of Biological Sciences, University of Southern California Los Angeles, Los Angeles, California, United States of America

David A. Caron

Affiliation Department of Biological Sciences, University of Southern California Los Angeles, Los Angeles, California, United States of America

Karla B. Heidelberg

kheidelb@usc.edu

Affiliation Department of Biological Sciences, University of Southern California Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AK ACJ DAC KBH. Performed the experiments: AK ACJ. Analyzed the data: AK ZL RT DAC KBH. Contributed reagents/materials/analysis tools: AK ZL RT DAC KBH. Wrote the paper: AK RT DAC KBH.