About the Authors

Rachel A. Levin

Contributed equally to this work with: Rachel A. Levin, Csilla N. Felsen

Affiliation Department of Pharmacology, UCSD School of Medicine, University of California San Diego, La Jolla, California, United States of America

Csilla N. Felsen

Contributed equally to this work with: Rachel A. Levin, Csilla N. Felsen

Affiliation Department of Pharmacology, UCSD School of Medicine, University of California San Diego, La Jolla, California, United States of America

Jin Yang

Affiliation Department of Pharmacology, UCSD School of Medicine, University of California San Diego, La Jolla, California, United States of America

John Y. Lin

Affiliation Department of Pharmacology, UCSD School of Medicine, University of California San Diego, La Jolla, California, United States of America

Michael A. Whitney

Affiliation Department of Pharmacology, UCSD School of Medicine, University of California San Diego, La Jolla, California, United States of America

Quyen T. Nguyen

Affiliation Division of Otolaryngology/Head and Neck Surgery, University of California San Diego, La Jolla, California, United States of America

Roger Y. Tsien

rtsien@ucsd.edu

Affiliations Department of Pharmacology, UCSD School of Medicine, University of California San Diego, La Jolla, California, United States of America, Howard Hughes Medical Institute, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RYT RAL CNF JY QTN. Performed the experiments: RAL CNF JY JYL MAW. Analyzed the data: RAL CNF JY. Contributed reagents/materials/analysis tools: RAL CNF JY JYL MAW. Wrote the paper: RAL CNF JY JYL MAW QTN RYT.