About the Authors

Masa Tsuchiya

tsuchiya@ttck.keio.ac.jp (MT); keyoshik@mail.doshisha.ac.jp (KY)

Affiliations Institute for Advanced Biosciences, Keio University, Tsuruoka, Japan, Systems Biology Program, School of Media and Governance, Keio University, Fujisawa, Japan

Midori Hashimoto

Affiliation Graduate School of Frontier Science, The University of Tokyo, Kashiwa, Japan

Yoshiko Takenaka

Affiliation Nanosystem Research Institute, National Institute of Advanced Industrial Science and Technology, Tsukuba, Japan

Ikuko N. Motoike

Affiliation Tohoku Medical Megabank Organization, Tohoku University, Sendai, Japan

Kenichi Yoshikawa

tsuchiya@ttck.keio.ac.jp (MT); keyoshik@mail.doshisha.ac.jp (KY)

Affiliation Faculty of Life and Medical Sciences, Doshisha University, Kyotanabe, Japan

Competing Interests

Masa Tsuchiya is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: MT MH YT IM KY. Analyzed the data: MT MH KY. Wrote the paper: MT KY.