About the Authors

Lai He

Contributed equally to this work with: Lai He, Jin Zhao, Maggie Haitian Wang

Affiliation Stanley Ho Center for Emerging Infectious Diseases, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China

Jin Zhao

Contributed equally to this work with: Lai He, Jin Zhao, Maggie Haitian Wang

Affiliations Stanley Ho Center for Emerging Infectious Diseases, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China, Shenzhen Center for Disease Control and Prevention, Shenzhen, China

Maggie Haitian Wang

Contributed equally to this work with: Lai He, Jin Zhao, Maggie Haitian Wang

Affiliation Division of Biostatistics, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China

Kenny K. Y. Siu

Affiliation Division of Biosciences, Faculty of Life Science, University College London, London, United Kingdom

Yong-Xia Gan

Affiliation Shenzhen Center for Disease Control and Prevention, Shenzhen, China

Lin Chen

Affiliation Shenzhen Center for Disease Control and Prevention, Shenzhen, China

Benny C. Y. Zee

Affiliation Division of Biostatistics, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China

Li Yang

Affiliation Division of Digestive Diseases, West China Hospital, Sichuan University, Chengdu, China

Hsiang-Fu Kung

Affiliations Stanley Ho Center for Emerging Infectious Diseases, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China, Li Ka Shing Institute of Health Sciences, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China

Zheng-Rong Yang

zryang2012@gmail.com (ZRY); mlhe7788@gmail.com (MLH)

Affiliation Shenzhen Center for Disease Control and Prevention, Shenzhen, China

Ming-Liang He

zryang2012@gmail.com (ZRY); mlhe7788@gmail.com (MLH)

Affiliations Stanley Ho Center for Emerging Infectious Diseases, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China, Li Ka Shing Institute of Health Sciences, School of Public Health and Primary Care, the Chinese University of Hong Kong, Hong Kong SAR, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZRY MLH BZ. Performed the experiments: LH JZ YXG. Analyzed the data: LH JZ KS MHW. Contributed reagents/materials/analysis tools: JZ LY HFK LC. Wrote the paper: LH MHW MLH.