About the Authors

Callie A. Norris

Contributed equally to this work with: Callie A. Norris, Mu He

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Mu He

Contributed equally to this work with: Callie A. Norris, Mu He

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Liang-I Kang

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Michael Qi Ding

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Josiah E. Radder

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Meagan M. Haynes

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Yu Yang

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Shirish Paranjpe

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

William C. Bowen

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Anne Orr

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

George K. Michalopoulos

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Donna B. Stolz

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Wendy M. Mars

wmars@pitt.edu

Affiliation Department of Pathology, University of Pittsburgh, School of Medicine, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CAN MH L-IK JER GKM DBS WMM. Performed the experiments: CAN MH MQD MMH YY SP WCB AO DBS. Analyzed the data: CAN MH L-IK MQD DBS WMM. Contributed reagents/materials/analysis tools: GKM. Wrote the paper: CAN WMM.