About the Authors

Rui Huang

Affiliation Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Dana Faratian

Affiliation Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Andrew H. Sims

Affiliation Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Danielle Wilson

Affiliation Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Jeremy S. Thomas

Affiliation Department of Pathology, Western General Hospital, Edinburgh, Scotland, United Kingdom

David J. Harrison

Affiliation Pathology, Medical and Biological Sciences Building, University of St Andrews, North Haugh, St. Andrews, Fife, Scotland, United Kingdom

Simon P. Langdon

simon.langdon@ed.ac.uk

Affiliation Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RH DF AHS DW DJH SPL. Performed the experiments: RH DF AHS DW JST DJH SPL. Analyzed the data: RH DF AHS DW JST DJH SPL. Contributed reagents/materials/analysis tools: AHS JST. Wrote the paper: RH DF AHS DW JST DJH SPL.