About the Authors

Margaret J. Couvillon

M.Couvillon@Sussex.ac.uk

Affiliation Laboratory of Apiculture and Social Insects, School of Life Sciences, University of Sussex, Falmer, Brighton, United Kingdom

Roger Schürch

Affiliations Laboratory of Apiculture and Social Insects, School of Life Sciences, University of Sussex, Falmer, Brighton, United Kingdom, Laboratory of Social Evolution, School of Life Sciences, University of Sussex, Falmer, Brighton, United Kingdom

Francis L. W. Ratnieks

Affiliation Laboratory of Apiculture and Social Insects, School of Life Sciences, University of Sussex, Falmer, Brighton, United Kingdom

Competing Interests

The commercial funders Waitrose Ltd. and Burt's Bees supported research through a charitable donation that paid for equipment and to hire undergraduate research assistants. They were not involved in any aspect of the experimental design, analysis, and manuscript preparation. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MJC FLWR. Performed the experiments: MJC. Analyzed the data: MJC RS. Contributed reagents/materials/analysis tools: MJC RS. Wrote the paper: MJC. Commented on manuscript: MJC RS FLWR. Take full responsibility for its content: MJC RS FLWR.