About the Authors

Naira N. Johnston

johnstn@alumni.unbc.ca

Affiliation Ecosystem Science and Management, University of Northern British Columbia, Prince George, British Columbia, Canada

James E. Bradley

Affiliation Ecosystem Science and Management, University of Northern British Columbia, Prince George, British Columbia, Canada

Ken A. Otter

Affiliation Ecosystem Science and Management, University of Northern British Columbia, Prince George, British Columbia, Canada

Competing Interests

This study had a commercial funder, Stantec Consulting, as part of the requirements under the Natural Sciences and Research Engineering Council of Canada (NSERC) Industrial Postgraduate Scholarships program (http://www.nserc-crsng.gc.ca/students-etudiants/pg-cs/ips-besii_eng.asp). This, however, does not alter our adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NNJ JEB KAO. Performed the experiments: NNJ JEB. Analyzed the data: NNJ KAO. Contributed reagents/materials/analysis tools: NNJ JEB KAO. Wrote the paper: NNJ JEB KAO.