About the Authors

Elske N. de Haas

elske.dehaas@wur.nl

Affiliation Adaptation Physiology Group, Department of Animal Science, Wageningen University and Research, Wageningen, The Netherlands

J. Elizabeth Bolhuis

Affiliation Adaptation Physiology Group, Department of Animal Science, Wageningen University and Research, Wageningen, The Netherlands

Bas Kemp

Affiliation Adaptation Physiology Group, Department of Animal Science, Wageningen University and Research, Wageningen, The Netherlands

Ton G. G. Groothuis

Affiliation Behavioural Biology, Centre for Behaviour and Neuroscience, University of Groningen, Groningen, The Netherlands

T. Bas Rodenburg

Affiliation Behavioural Ecology Group, Department of Animal Sciences, Wageningen University and Research, Wageningen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EDH JEB TBR TGG BK. Performed the experiments: EDH TBR. Analyzed the data: EDH TBR JEB BK. Contributed reagents/materials/analysis tools: JEB BK. Wrote the paper: EDH JEB BK TGG TBR. Grant application: TBR.