About the Authors

William A. Beltran

Contributed equally to this work with: William A. Beltran, Artur V. Cideciyan

wbeltran@vet.upenn.edu (WAB); cideciya@mail.med.upenn.edu (AVC)

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Artur V. Cideciyan

Contributed equally to this work with: William A. Beltran, Artur V. Cideciyan

wbeltran@vet.upenn.edu (WAB); cideciya@mail.med.upenn.edu (AVC)

Affiliation Department of Ophthalmology, Scheie Eye Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Karina E. Guziewicz

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Simone Iwabe

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Malgorzata Swider

Affiliation Department of Ophthalmology, Scheie Eye Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Erin M. Scott

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Svetlana V. Savina

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gordon Ruthel

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Frank Stefano

Affiliation Department of Biochemistry, School of Dental Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Lingli Zhang

Affiliation Department of Neuroscience, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Richard Zorger

Affiliation Department of Neuroscience, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alexander Sumaroka

Affiliation Department of Ophthalmology, Scheie Eye Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Samuel G. Jacobson

Affiliation Department of Ophthalmology, Scheie Eye Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gustavo D. Aguirre

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WAB AVC SGJ GDA. Performed the experiments: WAB AVC SI EMS SVS GR FS LZ SGJ GDA. Analyzed the data: WAB AVC KEG MS GR LZ RZ AS SGJ GDA. Wrote the paper: WAB AVC SGJ GDA.