About the Authors

Christine K. Ellis

Affiliations Animal Population Health Institute, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America, United States Department of Agriculture, Animal Plant and Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Randal S. Stahl

Affiliation United States Department of Agriculture, Animal Plant and Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Pauline Nol

Affiliation United States Department of Agriculture, Animal Plant and Health Inspection Service, Veterinary Services, Wildlife Livestock Disease Investigations Team, Fort Collins, Colorado, United States of America

W. Ray Waters

Affiliation United States Department of Agriculture, Agricultural Research Service, National Animal Disease Center, Ames, Iowa, United States of America

Mitchell V. Palmer

Affiliation United States Department of Agriculture, Agricultural Research Service, National Animal Disease Center, Ames, Iowa, United States of America

Jack C. Rhyan

Affiliation United States Department of Agriculture, Animal Plant and Health Inspection Service, Veterinary Services, Wildlife Livestock Disease Investigations Team, Fort Collins, Colorado, United States of America

Kurt C. VerCauteren

Affiliation United States Department of Agriculture, Animal Plant and Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Matthew McCollum

Affiliation United States Department of Agriculture, Animal Plant and Health Inspection Service, Veterinary Services, Wildlife Livestock Disease Investigations Team, Fort Collins, Colorado, United States of America

M. D. Salman

m.d.salman@colostate.edu

Affiliation Animal Population Health Institute, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CKE RSS PN WRW MVP JCR MM KCV MDS. Performed the experiments: CKE RSS PN WRW MVP. Analyzed the data: CKE RSS. Contributed reagents/materials/analysis tools: RSS PN WRW MVP JCR MM KCV MDS. Wrote the paper: CKE RSS PN. Editing and final manuscript approval: WRW MVP JCR MM KCV MDS.