About the Authors

Joanna M. Marshall

Affiliation Department of Microbial Infection and Immunity, Center for Microbial Interface Biology, The Ohio State University, Columbus, Ohio, United States of America

Alan D. Flechtner

Affiliation Department of Veterinary Biosciences, Comparative Pathology and Mouse Phenotyping Shared Resource, The Ohio State University, Columbus, Ohio, United States of America

Krista M. La Perle

Affiliation Department of Veterinary Biosciences, Comparative Pathology and Mouse Phenotyping Shared Resource, The Ohio State University, Columbus, Ohio, United States of America

John S. Gunn

gunn.43@osu.edu

Affiliation Department of Microbial Infection and Immunity, Center for Microbial Interface Biology, The Ohio State University, Columbus, Ohio, United States of America

Competing Interests

Co-author John S. Gunn is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JMM JSG ADF KML. Performed the experiments: JMM ADF. Analyzed the data: JMM JSG. Contributed reagents/materials/analysis tools: JMM JSG ADF KML. Wrote the paper: JMM JSG ADF KML.