About the Authors

Taiju Saito

Affiliations Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan, Laboratory of Aquaculture Genetics & Genomics, Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Japan, Laboratory of Reproductive Physiology, Research Institute of Fish Culture and Hydrobiology, South Bohemian Research Center of Aquaculture and Biodiversity of Hydrocenoses, Faculty of Fisheries and Protection of Waters, University of South Bohemia in Ceske Budejovice, Vodňany, Czech Republic

Martin Pšenička

Affiliation Laboratory of Reproductive Physiology, Research Institute of Fish Culture and Hydrobiology, South Bohemian Research Center of Aquaculture and Biodiversity of Hydrocenoses, Faculty of Fisheries and Protection of Waters, University of South Bohemia in Ceske Budejovice, Vodňany, Czech Republic

Rie Goto

Affiliation Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan

Shinji Adachi

Affiliation Laboratory of Aquaculture Biology, Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Japan

Kunio Inoue

Affiliation Department of Biology, Faculty of Science, Kobe University, Kobe, Japan

Katsutoshi Arai

Affiliation Laboratory of Aquaculture Genetics & Genomics, Faculty of Fisheries Sciences, Hokkaido University, Hakodate, Japan

Etsuro Yamaha

Affiliation Nanae Fresh Water Laboratory, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TS. Performed the experiments: TS MP RG. Analyzed the data: TS MP RG. Contributed reagents/materials/analysis tools: KI SA KA EY. Wrote the paper: TS.