About the Authors

Masaru K. Hojo

m.k.hojo@gmail.com

Current address: Department of Biology, Graduate School of Science, Kobe University, Kobe, Japan

Affiliations Faculty of Agriculture, University of the Ryukyus, Nishihara, Okinawa, Japan, Graduate School of Science and Technology, Kyoto Institute of Technology, Kyoto, Japan, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, United States of America

Ari Yamamoto

Affiliation Graduate School of Science and Technology, Kyoto Institute of Technology, Kyoto, Japan

Toshiharu Akino

Affiliation Graduate School of Science and Technology, Kyoto Institute of Technology, Kyoto, Japan

Kazuki Tsuji

Affiliations Faculty of Agriculture, University of the Ryukyus, Nishihara, Okinawa, Japan, The United Graduate School of Agricultural Sciences, Kagoshima University, Kagoshima, Japan

Ryohei Yamaoka

Affiliation Graduate School of Science and Technology, Kyoto Institute of Technology, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MKH RY. Performed the experiments: MKH AY. Analyzed the data: MKH TA KT. Wrote the paper: MKH TA KT RY.