About the Authors

Li-Guo Li

dinoliguo@gmail.com

Affiliations School of Earth Sciences and Resources, China University of Geosciences (Beijing), Beijing, China, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Da-Qing Li

Affiliation Gansu Geological Museum, Lanzhou, China

Hai-Lu You

Affiliation Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, P. R. China

Peter Dodson

Affiliation School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

Co-author professor Peter Dodson is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DQL HLY. Performed the experiments: LGL. Analyzed the data: LGL PD. Contributed reagents/materials/analysis tools: LGL HLY DQL PD. Wrote the paper: LGL PD. Edited the manuscript: PD HLY.