About the Authors

David Moretti

david.moretti@navy.mil

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Len Thomas

Affiliation Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St. Andrews, Scotland

Tiago Marques

Affiliation Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St. Andrews, Scotland

John Harwood

Affiliation Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St. Andrews, Scotland

Ashley Dilley

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Bert Neales

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Jessica Shaffer

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Elena McCarthy

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Leslie New

Affiliation U.S. Marine Mammal Commission, Bethesda, Maryland, United States of America

Susan Jarvis

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Ronald Morrissey

Affiliation Naval Undersea Warfare Center, Newport, Rhode Island, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DM LT TM JH LN AD EM. Performed the experiments: DM BN JS SJ RM EM AD. Analyzed the data: DM LT TM JH AD EM JS BN. Contributed reagents/materials/analysis tools: DM TM LT SJ RM JS. Wrote the paper: DM TM LT JH AD LN BN JS EM RM SJ.