About the Authors

Yong-Ju Huang

y.huang8@herts.ac.uk

Affiliations School of Life and Medical Sciences, University of Hertfordshire, Hatfield, Hertfordshire, United Kingdom, Department of Plant Pathology and Microbiology, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom

Aiming Qi

Affiliations School of Life and Medical Sciences, University of Hertfordshire, Hatfield, Hertfordshire, United Kingdom, Department of Plant Pathology and Microbiology, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom

Graham J. King

Affiliations Department of Plant Pathology and Microbiology, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom, Southern Cross Plant Science, Southern Cross University, Lismore, Australia

Bruce D. L. Fitt

Affiliations School of Life and Medical Sciences, University of Hertfordshire, Hatfield, Hertfordshire, United Kingdom, Department of Plant Pathology and Microbiology, Rothamsted Research, Harpenden, Hertfordshire, United Kingdom

Competing Interests

The authors have the following interests: This study was partly funded by DuPont. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: YJH GJK BDLF. Performed the experiments: YJH. Analyzed the data: YJH AQ. Contributed reagents/materials/analysis tools: YJH AQ GJK. Wrote the paper: YJH AQ BDLF.