About the Authors

Chin-Yi Chen

Contributed equally to this work with: Chin-Yi Chen, Gaylen A. Uhlich

chin-yi.chen@ars.usda.gov

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Christopher S. Hofmann

Current address: United States Army Medical Research Institute of Chemical Defense, Aberdeen, Maryland, United States of America

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Bryan J. Cottrell

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Terence P. Strobaugh Jr

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

George C. Paoli

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Ly-Huong Nguyen

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Xianghe Yan

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Gaylen A. Uhlich

Contributed equally to this work with: Chin-Yi Chen, Gaylen A. Uhlich

Affiliation Molecular Characterization of Foodborne Pathogens Research Unit, Eastern Regional Research Center, Agricultural Research Service, United States Department of Agriculture, Wyndmoor, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CYC GAU. Performed the experiments: CYC CSH BJC TPS GCP LHN GAU. Analyzed the data: CYC CSH BJC TPS GCP LHN XY GAU. Contributed reagents/materials/analysis tools: GCP XY. Wrote the manuscript: CYC GAU. Drafted and edited the paper: CYC CSH BJC TPS GCP LHN XY GAU.