About the Authors

Mariel M. Finucane

Affiliation The J. David Gladstone Institutes, San Francisco, California, United States of America

Thomas J. Sharpton

Current address: Departments of Microbiology and Statistics, Oregon State University, Corvallis, Oregon, United States of America

Affiliation The J. David Gladstone Institutes, San Francisco, California, United States of America

Timothy J. Laurent

Affiliation The J. David Gladstone Institutes, San Francisco, California, United States of America

Katherine S. Pollard

kpollard@gladstone.ucsf.edu

Affiliations The J. David Gladstone Institutes, San Francisco, California, United States of America, Institute for Human Genetics and Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MMF TJS KSP. Analyzed the data: MMF TJS TL. Wrote the paper: MMF TJS TL KSP.