About the Authors

Jean A. Hall

Jean.Hall@oregonstate.edu

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

William R. Vorachek

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

Whitney C. Stewart

Affiliations Department of Animal and Rangeland Sciences, College of Agriculture, Oregon State University, Corvallis, Oregon, United States of America, Current Address: Department of Animal and Range Sciences, New Mexico State University, Las Cruces, New Mexico, United States of America

M. Elena Gorman

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Oregon State University, Corvallis, Oregon, United States of America

Wayne D. Mosher

Affiliation Department of Animal and Rangeland Sciences, College of Agriculture, Oregon State University, Corvallis, Oregon, United States of America

Gene J. Pirelli

Affiliation Department of Animal and Rangeland Sciences, College of Agriculture, Oregon State University, Corvallis, Oregon, United States of America

Gerd Bobe

Affiliations Department of Animal and Rangeland Sciences, College of Agriculture, Oregon State University, Corvallis, Oregon, United States of America, Linus Pauling Institute, Oregon State University, Corvallis, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAH WDM GJP . Performed the experiments: JAH WRV WCS MEG WDM GJP . Analyzed the data: JAH GB. Wrote the paper: JAH GB.