About the Authors

Erich A. Heise

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Lauren M. Marozas

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Sean A. Grafton

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Katelyn M. Green

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Stefanie J. Kirwin

Current address: Eye.Q.Consulting, Irvine, California, United States of America

Affiliation Biological Science, Allergan Incorporated, Irvine, California, United States of America

Patrice E. Fort

patricef@umich.edu

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

Stefanie J. Kirwin is a consultant for Allergan, Inc., but this does not alter the authors' adherence to all the PLOS ONE policies on sharting data and materials.

Author Contributions

Conceived and designed the experiments: SJK PEF. Performed the experiments: EAH LMM SAG KMG PEF. Analyzed the data: SAG KMG PEF. Contributed reagents/materials/analysis tools: SJK PEF. Wrote the paper: EAH SJK PEF.