About the Authors

Florent Bled

florent.bled@gmail.com

Affiliations US Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America, Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America

John Sauer

Affiliation US Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Keith Pardieck

Affiliation US Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Paul Doherty

Affiliation Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America

J. Andrew Royle

Affiliation US Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JS KP. Performed the experiments: FB JS KP JAR. Analyzed the data: FB JAR JS. Contributed reagents/materials/analysis tools: FB JAR JS. Wrote the paper: FB JAR JS PD KP.