About the Authors

Lucy A. Taylor

ltaylor@bristolzoo.org.uk

Affiliations Bristol Conservation and Science Foundation, c/o Bristol Zoo Gardens, Bristol, United Kingdom, School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Thomas M. Kaiser

Affiliation Biocenter Grindel and Zoological Museum, University of Hamburg, Hamburg, Germany

Christoph Schwitzer

Affiliation Bristol Conservation and Science Foundation, c/o Bristol Zoo Gardens, Bristol, United Kingdom

Dennis W. H. Müller

Affiliations Clinic for Zoo Animals, Exotic Pets and Wildlife, Vetsuisse Faculty, University of Zurich, Zurich, Switzerland, National Park ‘Bavarian Forest’, Grafenau, Germany

Daryl Codron

Affiliations Clinic for Zoo Animals, Exotic Pets and Wildlife, Vetsuisse Faculty, University of Zurich, Zurich, Switzerland, Florisbad Quaternary Research Department, National Museum, Bloemfontein, South Africa

Marcus Clauss

Affiliation Clinic for Zoo Animals, Exotic Pets and Wildlife, Vetsuisse Faculty, University of Zurich, Zurich, Switzerland

Ellen Schulz

Affiliation Biocenter Grindel and Zoological Museum, University of Hamburg, Hamburg, Germany

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK MC ES LT. Performed the experiments: LT ES. Analyzed the data: LT DC ES MC. Contributed reagents/materials/analysis tools: DM ES TK MC. Wrote the paper: LT ES TK MC CS.