About the Authors

Jing Yang

Contributed equally to this work with: Jing Yang, Hui Yu

Affiliations School of Biotechnology, East China University of Science and Technology, Shanghai, P. R. China, Bioinformatics Center, Key Laboratory of Systems Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P. R. China

Hui Yu

Contributed equally to this work with: Jing Yang, Hui Yu

Affiliations Shanghai Center for Bioinformation Technology, Shanghai Industrial Technology Institute, Shanghai, P. R. China, Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Bao-Hong Liu

Affiliation Shanghai Center for Bioinformation Technology, Shanghai Industrial Technology Institute, Shanghai, P. R. China

Zhongming Zhao

Affiliations Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Departments of Psychiatry, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Lei Liu

Affiliation Bioinformatics Center, Key Laboratory of Systems Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P. R. China

Liang-Xiao Ma

Affiliation Shanghai Center for Bioinformation Technology, Shanghai Industrial Technology Institute, Shanghai, P. R. China

Yi-Xue Li

yyli@scbit.org (YYL); yxli@scbit.org (YXL)

Affiliations School of Biotechnology, East China University of Science and Technology, Shanghai, P. R. China, Shanghai Center for Bioinformation Technology, Shanghai Industrial Technology Institute, Shanghai, P. R. China

Yuan-Yuan Li

yyli@scbit.org (YYL); yxli@scbit.org (YXL)

Affiliation Shanghai Center for Bioinformation Technology, Shanghai Industrial Technology Institute, Shanghai, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YYL HY JY YXL. Performed the experiments: JY HY BHL. Analyzed the data: JY HY. Contributed reagents/materials/analysis tools: YYL JY HY LXM LL. Wrote the paper: YYL HY JY ZZ.