About the Authors

Sara J. Elliott

Affiliation Department of Geosciences, Pennsylvania State University, University Park, Pennsylvania, United States of America

Peter Wilf

pwilf@psu.edu

Affiliation Department of Geosciences, Pennsylvania State University, University Park, Pennsylvania, United States of America

Robert C. Walter

Affiliation Department of Earth and Environment, Franklin and Marshall College, Lancaster, Pennsylvania, United States of America

Dorothy J. Merritts

Affiliation Department of Earth and Environment, Franklin and Marshall College, Lancaster, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJE PW. Performed the experiments: SJE. Analyzed the data: SJE PW. Contributed reagents/materials/analysis tools: SJE PW. Wrote the paper: SJE. Discovered fossil site and contributed substantially to data collection and interpretation: RCW DJM. Commented on and substantially edited manuscript: PW RCW DJM.