About the Authors

Damian Madan

damianmadan@gmail.com (DM); ctesta@echelon-inc.com (CT)

Current address: Intellectual Ventures Laboratories, Seattle, Washington, United States of America

Affiliation Echelon Biosciences Inc., Salt Lake City, Utah, United States of America

Colin G. Ferguson

Affiliation Echelon Biosciences Inc., Salt Lake City, Utah, United States of America

Won Yong Lee

Affiliation Department of Medicinal Chemistry, University of Utah, Salt Lake City, Utah, United States of America

Glenn D. Prestwich

Affiliation Department of Medicinal Chemistry, University of Utah, Salt Lake City, Utah, United States of America

Charles A. Testa

damianmadan@gmail.com (DM); ctesta@echelon-inc.com (CT)

Affiliation Echelon Biosciences Inc., Salt Lake City, Utah, United States of America

Competing Interests

C. Ferguson and C. Testa are currently employed, D. Madan was previously employed, and G.D. Prestwich is a paid consultant of Echelon Biosciences. No compensation was linked to the use of these research tools. The imaging probe (AR-2) described in this manuscript is currently marketed by Echelon Biosciences under catalog number L-2010 and is covered under U.S. patent number 8,512,971. These factors do not alter the authors' adherence to the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DM CF WYL CT GP. Performed the experiments: DM CF WYL. Analyzed the data: DM CF WYL. Wrote the paper: DM CT.