About the Authors

Joshua D. Ramsay

jdr105@vetmed.wsu.edu

Affiliation Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America

Massaro W. Ueti

Affiliation Animal Disease Research Unit, Agricultural Research Service, USDA, Pullman, Washington, United States of America

Wendell C. Johnson

Affiliation Animal Disease Research Unit, Agricultural Research Service, USDA, Pullman, Washington, United States of America

Glen A. Scoles

Affiliation Animal Disease Research Unit, Agricultural Research Service, USDA, Pullman, Washington, United States of America

Donald P. Knowles

Affiliations Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America, Animal Disease Research Unit, Agricultural Research Service, USDA, Pullman, Washington, United States of America

Robert H. Mealey

Affiliation Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDR MWU WCJ GAS DPK RHM. Performed the experiments: JDR MWU WCJ. Analyzed the data: JDR MWU WCJ GAS DPK RHM. Contributed reagents/materials/analysis tools: JDR MWU GAS DPK RHM. Wrote the paper: JDR RHM.