About the Authors

Anne Marijke Schel

Affiliations Department of Psychology, University of York, York, United Kingdom, Budongo Conservation Field Station, Masindi, Uganda

Simon W. Townsend

Affiliation Animal Behaviour Group, Institute of Evolutionary Biology and Environmental Studies, University of Zurich, Zurich, Switzerland

Zarin Machanda

Affiliation Department of Human Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Klaus Zuberbühler

Affiliations Budongo Conservation Field Station, Masindi, Uganda, Department of Comparative Cognition, University of Neuchâtel, Neuchâtel, Switzerland, School of Psychology & Neurosciences, University of St Andrews, St Andrews, United Kingdom

Katie E. Slocombe

katie.slocombe@york.ac.uk

Affiliation Department of Psychology, University of York, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KS ST AS KZ. Performed the experiments: KS AS. Analyzed the data: KS AS ST ZM. Wrote the paper: KS AS ST ZM KZ.