About the Authors

Elizabeth D. Tobin

etobin@u.washington.edu

Affiliation School of Oceanography, University of Washington, Seattle, Washington, United States of America

Daniel Grünbaum

Affiliation School of Oceanography, University of Washington, Seattle, Washington, United States of America

Johnathan Patterson

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Rose Ann Cattolico

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ET DG RAC. Performed the experiments: ET JP. Analyzed the data: ET JP. Contributed reagents/materials/analysis tools: ET DG RAC. Wrote the manuscript: ET DG RAC. Manuscript edits and revisions: ET DG RAC JP. Designed video-based particle tracking software used in this analysis: DG.