About the Authors

Bruce A. Kimball

bruce.a.kimball@aphis.usda.gov

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Philadelphia, Pennsylvania, United States of America

Kunio Yamazaki

† Deceased.

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Dennis Kohler

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Richard A. Bowen

Affiliation Department of Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Jack P. Muth

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Maryanne Opiekun

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Gary K. Beauchamp

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BAK KY RAB GKB. Performed the experiments: KY DK JPM MO. Analyzed the data: BAK KY GKB. Contributed reagents/materials/analysis tools: DK RAB JPM. Wrote the paper: BAK DK GKB.