About the Authors

Bianca Haase

bianca.haase@sydney.edu.au

Affiliations Faculty of Veterinary Science, University of Sydney, Sydney, New South Wales, Australia, Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Heidi Signer-Hasler

Affiliation Department of Agriculture, Forestry, Food Science and Management, Bern University of Applied Science, Zollikofen, Switzerland

Matthew M. Binns

Affiliation Equine Analysis, Midway, Kentucky, United States of America

Gabriela Obexer-Ruff

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Regula Hauswirth

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Rebecca R. Bellone

Affiliation Department of Biology, University of Tampa, Tampa, Florida, United States of America

Dominik Burger

Affiliation Swiss Institute of Equine Medicine, ALP-Haras and University of Bern, Avenches, Switzerland

Stefan Rieder

Contributed equally to this work with: Stefan Rieder, Claire M. Wade, Tosso Leeb

Affiliation Agroscope ALP-Haras Research Station, Swiss National Stud Farm, Avenches, Switzerland

Claire M. Wade

Contributed equally to this work with: Stefan Rieder, Claire M. Wade, Tosso Leeb

Affiliation Faculty of Veterinary Science, University of Sydney, Sydney, New South Wales, Australia

Tosso Leeb

Contributed equally to this work with: Stefan Rieder, Claire M. Wade, Tosso Leeb

Affiliations Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland, DermFocus, University of Bern, Bern, Switzerland

Competing Interests

The co-author Matthew M. Binns is affiliated with Equine Analysis. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: BH SR CMW TL. Performed the experiments: BH. Analyzed the data: BH HSH MMB RH RRB. Contributed reagents/materials/analysis tools: GOR DB SR. Wrote the paper: BH CMW.