About the Authors

Andrew J. Elmore

aelmore@umces.edu

Affiliation University of Maryland Center for Environmental Science, Appalachian Laboratory, Frostburg, Maryland, United States of America

Jason P. Julian

Affiliation Department of Geography, Texas State University, San Marcos, Texas, United States of America

Steven M. Guinn

Affiliation University of Maryland Center for Environmental Science, Appalachian Laboratory, Frostburg, Maryland, United States of America

Matthew C. Fitzpatrick

Affiliation University of Maryland Center for Environmental Science, Appalachian Laboratory, Frostburg, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AJE JPJ. Performed the experiments: AJE JPJ SMG MCF. Analyzed the data: AJE SMG MCF. Contributed reagents/materials/analysis tools: SMG MCF. Wrote the manuscript: AJE JPJ MCF.