About the Authors

Haifa Benhajali

Affiliations UMR CNRS 6552 Ethologie Animale et Humaine, Université de Rennes I, Rennes, France, Haras national de Sidi Thabet, Sidi Thabet, Tunisia

Mohammed Ezzaouia

Affiliation Haras national de Sidi Thabet, Sidi Thabet, Tunisia

Christophe Lunel

Affiliation UMR CNRS 6552 Ethologie Animale et Humaine, Université de Rennes I, Rennes, France

Faouzia Charfi

Affiliation Unité de Biologie Animale et de Systématique Evolutive, Université Tunis-ElManar, Campus universitaire, Tunis, Tunisia

Martine Hausberger

martine.hausberger@univ-rennes1.fr

Affiliation UMR CNRS 6552 Ethologie Animale et Humaine, Université de Rennes I, Rennes, France

Competing Interests

The “Haras National of Sidi Thabet” made its horses available for the present study but had no role in study design, data collection and analysis, decision to publish or preparation of the manuscript. Dr Ezzaouia was affiliated to this company. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HB MH. Performed the experiments: HB ME FC. Analyzed the data: HB CL MH. Wrote the paper: HB MH.