About the Authors

Darren Pais

Affiliation Department of Mechanical and Aerospace Engineering, Princeton University, Princeton, New Jersey, United States of America

Patrick M. Hogan

Affiliation Department of Computer Science and Kroto Research Institute, University of Sheffield, Sheffield, South Yorkshire, United Kingdom

Thomas Schlegel

Affiliation School of Biological Sciences, University of Bristol, Bristol, Bristol, United Kingdom

Nigel R. Franks

Affiliation School of Biological Sciences, University of Bristol, Bristol, Bristol, United Kingdom

Naomi E. Leonard

Affiliation Department of Mechanical and Aerospace Engineering, Princeton University, Princeton, New Jersey, United States of America

James A. R. Marshall

James.Marshall@sheffield.ac.uk

Affiliation Department of Computer Science and Kroto Research Institute, University of Sheffield, Sheffield, South Yorkshire, United Kingdom

Competing Interests

The authors note that co-author JARM is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DP PMH NEL JARM. Performed the experiments: DP. Analyzed the data: DP PMH NEL JARM. Contributed reagents/materials/analysis tools: DP PMH JARM. Wrote the paper: DP PMH TS NRF NEL JARM.