About the Authors

Erika S. Wittchen

Contributed equally to this work with: Erika S. Wittchen, Eiichi Nishimura

Affiliation Department of Cell Biology and Physiology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Eiichi Nishimura

Contributed equally to this work with: Erika S. Wittchen, Eiichi Nishimura

Affiliations Moran Eye Center, University of Utah, Salt Lake City, Utah, United States of America, Department of Ophthalmology, Showa University, Tokyo, Japan

Manabu McCloskey

Affiliation Moran Eye Center, University of Utah, Salt Lake City, Utah, United States of America

Haibo Wang

Affiliation Moran Eye Center, University of Utah, Salt Lake City, Utah, United States of America

Lawrence A. Quilliam

Affiliation Department of Biochemistry and Molecular Biology, Indiana University, Indianapolis, Indiana, United States of America

Magdalena Chrzanowska-Wodnicka

Affiliation Blood Research Institute, Blood Center of Wisconsin, Milwaukee, Wisconsin, United States of America

M. Elizabeth Hartnett

ME.Hartnett@hsc.utah.edu

Affiliation Moran Eye Center, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors note here that M. Elizabeth Hartnett is an Academic Editor for PLOS ONE. However, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ESW MEH. Performed the experiments: ESW EN MM HW. Analyzed the data: ESW EN MM HW MEH. Contributed reagents/materials/analysis tools: LAQ MCW. Wrote the paper: ESW EN MM MEH.