About the Authors

Samantha L. Eaton

Affiliation Division of Neurobiology, The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Sarah L. Roche

Affiliation Centre for Integrative Physiology, University of Edinburgh, Edinburgh, United Kingdom

Maica Llavero Hurtado

Affiliation Division of Neurobiology, The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Karla J. Oldknow

Affiliation Developmental Biology, The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Colin Farquharson

Affiliation Developmental Biology, The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Thomas H. Gillingwater

Affiliations Centre for Integrative Physiology, University of Edinburgh, Edinburgh, United Kingdom, Euan MacDonald Centre for Motor Neurone Disease Research, University of Edinburgh, Edinburgh, United Kingdom

Thomas M. Wishart

T.M.Wishart@Roslin.ed.ac.uk

Affiliations Division of Neurobiology, The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom, Euan MacDonald Centre for Motor Neurone Disease Research, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors can confirm that author THG is currently an academic editor at PLOS ONE. This does not alter the authors’ adherence to all of the PLOS ONE

Author Contributions

Conceived and designed the experiments: TMW SLE. Performed the experiments: SLE SLR ML KO. Analyzed the data: SLE SLR ML KO CF THG TMW. Contributed reagents/materials/analysis tools: CF THG TMW. Wrote the paper: SLE THG TMW.