About the Authors

Mona J. Nolde

Affiliations Yale Stem Cell Center, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Cell Biology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Ee-chun Cheng

Affiliations Yale Stem Cell Center, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Cell Biology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Shangqin Guo

Affiliations Yale Stem Cell Center, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America

Haifan Lin

haifan.lin@yale.edu

Affiliations Yale Stem Cell Center, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Cell Biology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Competing Interests

The authors declare no competing interests.

Author Contributions

Conceived and designed the experiments: MN EC HL. Performed the experiments: MN EC SG. Analyzed the data: MN EC SG HL. Contributed reagents/materials/analysis tools: EC SG. Wrote the paper: MN HL EC.