About the Authors

Jie Lv

Contributed equally to this work with: Jie Lv, Wei Cui, Hongbo Liu

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Wei Cui

Contributed equally to this work with: Jie Lv, Wei Cui, Hongbo Liu

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Hongbo Liu

Contributed equally to this work with: Jie Lv, Wei Cui, Hongbo Liu

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Hongjuan He

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Youcheng Xiu

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Jing Guo

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Hui Liu

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Qi Liu

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Tiebo Zeng

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Yan Chen

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Yan Zhang

Affiliation College of Bioinformatics Science and Technology, Harbin Medical University, Harbin, China

Qiong Wu

kigo@hit.edu.cn

Affiliation School of Life Science and Technology, State Key Laboratory of Urban Water Resource and Environment, Harbin Institute of Technology, Harbin, China

Competing Interests

Qiong Wu is currently an academic editor. This does not alter the authors′ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YZ QW. Performed the experiments: JL WC HBL. Analyzed the data: HH YX JG. Contributed reagents/materials/analysis tools: HL QL TZ YC. Wrote the paper: JL WC HBL.