About the Authors

J. Jeffrey Root

Jeff.root@aphis.usda.gov

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Susan A. Shriner

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Kevin T. Bentler

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Thomas Gidlewski

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Disease Program, Fort Collins, Colorado, United States of America

Nicole L. Mooers

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Jeremy W. Ellis

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Terry R. Spraker

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Kaci K. VanDalen

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Heather J. Sullivan

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Alan B. Franklin

Affiliation United States Department of Agriculture, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJR SAS KKV ABF. Performed the experiments: JJR SAS KTB NLM TG JE. Analyzed the data: JJR SAS KTB TG NLM TRS HJS. Contributed reagents/materials/analysis tools: TG TRS. Wrote the paper: JJR SAS KTB TG KKV ABF.